Trees, shrubs or herbs, succulent or not, monoecious. Exudate present, white. Armed (sometimes with long, rigid stinging hairs) or unarmed. Indumentum present, hairs usually stinging, sometimes long, or hairs simple*. Stipules present*, free*, simple, unlobed, entire *, not spine-like*, not leaf-like*, without glands, or stipules 0*. Leaves distributed evenly along stem, alternate, simple or compound, lobed or unlobed, base symmetrical, margin toothed or entire, blade not pellucid-punctate, stipels at leaflet base present or absent *, stipel-like structures or glands present on lower surface at leaf base or petiole apex*; with 3 – 7 strong veins from base or venation pinnate; domatia absent*. Petiole present, without swellings or bends at base or apex*, with or without glands or stipel-like structures at apex. Inflorescence with axis, axillary or terminal, panicle-like; bracts not leaf-like*, green. Male flowers: pedicels present, articulated, or pedicels 0*; sepals 5; imbricate, partially fused, petal-like, greenish or white; petals 0; disc glands 5, extrastaminal*, stamens 10–20, free to partially or completely fused, in 1–more rows; anther with or without conspicuous modification, thecae introrse in bud; staminodes sometimes present at apex of stamen column; pistillode absent. Female flowers: pedicels present, not articulated, or pedicels 0*; sepals 5; imbricate, partially fused, reflexed, falling early, greenish or white; petals 0; disc glands 5, surrounding the ovary, ovary locules 3, ovules per locule 1; styles 3, partially fused, style 2–more-divided, (i.e. ultimate divisions per locule 2–many), styles or stigmas not flat*, surface smooth*; staminodes absent. Fruit dehiscent, not sculptured*, hairy, greenish to brown; seed with fleshy structure (aril? caruncle?) at one end; embryo flat, not minute, cotyledons leaf-like*.
An asterisk * indicates a state has been assumed but not yet confirmed.