Tree or shrub, dioecious. Exudate absent*. Unarmed*. Indumentum present, hairs simple. Stipules free*, simple, unlobed, entire *, not spine-like*, not leaf-like*, without glands. Leaves distributed evenly along stem, alternate, simple, unlobed, base symmetrical, margin entire, leaf blade not pellucid-punctate; leaf glands absent*; venation pinnate; domatia absent*; stipel-like appendages absent*. Petiole present, without swellings or bends at base or apex*, without glands. Inflorescence with axis, axillary, the males elongate, spike-like or paniculate, the females shorter, spicate to racemose; bracts not leaf-like*, green; pedicels present, not articulated*. Male flowers: pedicels short; sepals 4, imbricate, partially fused, greenish; petals 0; disc inside the staminal whorl (intrastaminal), cushion- or plate-shaped, 4-lobed; stamens 4, free; anther without conspicuous modification, thecae extrorse in bud; pistillode present. Female flowers: pedicels short, extending and thickening in fruit; sepals 5 – 6, imbricate, partially fused; greenish; petals 0; disc annular, surrounding the ovary; ovary locules 3, ovules per locule 2; styles 3, free* or partially fused*, very short, entire, styles or stigmas not flat*, surface smooth*; staminodes absent. Fruit dehiscent, unsculptured*, hairy, greenish to brown; seed without appendage, covering or membraneous layer*, not fleshy; embryo flat, not minute, cotyledons leaf-like.
An asterisk * indicates a state has been assumed but not yet confirmed.