Tree, dioecious. Exudate absent*. Unarmed*. Indumentum present, hairs simple. Stipules free*, simple, unlobed, entire *, not spine-like*, not leaf-like*, without glands. Leaves distributed evenly along stem, alternate, compound; leaflets unlobed, base symmetrical, margin toothed or entire, blade not pellucid-punctate, glands absent, venation pinnate, domatia absent*; stipel-like appendages absent*. Petiole present, without swellings or bends at base or apex*, without glands. Inflorescences axillary, often arising from the axis of fallen leaves, male inflorescences with axis, slender, raceme-like, female inflorescence with or without axis, racemose or flowers solitary; bracts present, not leaf-like*, green, or bracts 0; pedicels present, not articulated*. Male flowers: sepals 4 – 6, imbricate, free, unequal, greenish; petals 0; disc glands interstaminal, randomly distributed among stamens; stamens 8 – 16, free; anther papillose, otherwise without conspicuous modification, thecae in bud introrse, extrorse or latrorse; pistillode absent. Female flowers: pedicels often long, sepals 6 – 9, imbricate, free or partially fused, unequal, greenish, sometimes falling; petals 0; disc surrounding the ovary, annular, shallowly lobed; ovary locules 3, ovules per locule 2; styles (1) 3 (6), free to completely fused (stigmas may be free), styles 2-branched, (i.e. each style divided, ultimate divisions per locule 2), styles or stigmas not flat*, surface papillose; staminodes present or absent. Fruit dehiscent, unsculptured*, greenish to brown; seed without caruncle (or almost so), without covering or membraneous layer*, not fleshy;embryo flat, not minute, cotyledons leaf-like.
An asterisk * indicates a state has been assumed but not yet confirmed.